Table 2: Comparative interspecific profiles of scent marking behaviours among lemurid species of the genera Eulemur and Hapalemur, as derived from the published literature on these taxa [13, 90113]. The patterns seen in the two pair-bonded Eulemur species, E. mongoz and E. rubriventer, are very similar to each other, and contrast in significant ways to the patterns exhibited amongst the Eulemur taxa that occur in multimale, multifemale social groups (i.e., the E. fulvus group of taxa, E. macaco, and E. coronatus).

Scent-marking behavioursE. fulvus groupE. macacoE. coronatusE. mongozE. rubriventerHapalemur

Anogenital scent-mark surfaces++++++
Urine mark++++?+
Bite mark (saliva deposition) males+++Cheek rubbingTeeth grinding during inter-group encounters+
Anogenital mark Conspecifics M F M F M FM FM F M F
F MF M
HeadMarking (males)+++++
Hand/palmarmarking (males)+++++Antebrachial gland marking
Anogenital sniff (M F)++++++
Self-markingObserved in captivityObserved in captivityM marks tail with antebrachial glands to “stink fight” other Ms
Marking during inter-group encountersVariable.Vocal & visual displays; some markingVocal and visual displaysVocal & visual displays+++
(Incl. F overmarking of M anogen marks)(Both substrate and self-marking)
Flehmen?+????
(pers. obs.)
Latrine behaviour+