Research Article

Integrative Taxonomy, Phylogeny, and New Species of the Weevil Genus Onyxacalles Stüben (Coleoptera: Curculionidae: Cryptorhynchinae)

Table 1

Collecting data, vouchers, and GenBank accession numbers for the material analysed in this study. All specimens determined by P. E. Stüben, 2010 and 2011. Vouchers (DNA, morphology) are kept at the ZFMK Biobank. Most sequences have been published (or are reviewed) in Astrin et al. 2012 [21]. GenBank accession numbers of new sequences in this study start with “JN…”. Taxonomic changes with regard to this publication are printed in brackets (old name).

TaxonCollecting dataDNA voucherCOI 16S

Acalles camelus (Fabricius 1792)France: Isère, 2 km SE Lans en Vercors, Montagne de Lans; N45°06′45′′ E05°36′21′′, 1352 m; Abies, Fagus, Fraxinus, 2005, StübenZFMK-DNA-JJ0017, ZFMK-TIS-cI0026camEU286282 EU286447
Acallorneuma doderoi A. and F. Solari 1908Italy: Sicilia (PA), 6 km SW Godrano, Bosco Ficuzza, Mte. Rocca Busambra; N37°51′38′′ E13°23′24′′, 1200 m; Quercus, Fraxinus, 2002, StübenZFMK-DNA-JJ0065, ZFMK-TIS-cS0082dodEU286292 EU286457
Cryptorhynchus lapathi (Linné 1758)Germany: Bienen bei Rees, Altrheinarm, 2004, ScharfZFMK-DNA-JJ0214, ZFMK-TIS-cD0354lapEU286360 EU286523
Kyklioacalles fausti (Meyer 1896) [25]Italy: Campania, Cilento, 6 km SE Vallo d. Lucania, M. Sacro o Gelbison; N40°12′41′′ E15°19′42′′, 1544 m; Fagus, 2008, StübenZFMK-DNA-JJ0564, ZFMK-TIS-cI625fauGU213776 GU213772
Kyklioacalles pyrenaeus (Boheman 1844) [26] (gen. Onyxacalles)France: Isère, 14 km N Grenoble, Massif de la Chartreuse, NW Col de Porte; N45°18′40′′ E05°45′17′′, 1649 m; Abies, Fagus, Fraxinus, 2005, StübenZFMK-DNA-JJ0026, ZFMK-TIS-cI0035pyrGU988172 GU987762
Kyklioacalles pyrenaeus [26] (gen. Onyxacalles)Czech Republic: W Bohemia (KT), Balkovy, Doubrava Hill (6545), 2008, KreslZFMK-DNA-JJ0764, ZFMK-TIS-cCz798pyrGU981555 GU981506
Kyklioacalles reginae Stüben 2003Spain: Teruel, S. Javalambre, Fuente la Risca near Arcos de las Salinas; N39°59′56′′ W01°01′21′′, 1121 m; Amelanchier ovalis, Acer monspessulanum, Erinacea anthyllis, Ulex, 2008, StübenZFMK-DNA-JJ0539, ZFMK-TIS-cE600regGU981544 GU981495
Onyxacalles balearicus Stüben 2005Spain: Mallorca, 3 km SE Lluc, Sra. de Tramuntana, Sa Maleta; N39°48′47′′ E02°53′23′′, 571 m; Quercus ilex, 2004, StübenZFMK-DNA-JJ0207, ZFMK-TIS-cE0168balEU286357 EU286521
Onyxacalles balearicus Stüben 2005Spain: Mallorca, 11 km NE Lluc, Sra. de Tramuntana; N39°52′03′′ E02°58′20′′, 107 m; PT, Smilax aspera, Quercus ilex, 2004, StübenZFMK-DNA-JJ0466, ZFMK-TIS-cE0294balGU988348 _________
Onyxacalles bermejaensis Stüben 2001Spain: Andalucía, 11 km S Ronda, Sierra de las Nieves; N36°39′51′′ W05°05′01′′, 1047 m; Quercus ilex, 2005, StübenZFMK-DNA-JJ0198, ZFMK-TIS-cE0167berEU286350 EU286514
Onyxacalles bermejaensis Stüben 2001Spain: Málaga, 9 km SE Ubrique, Sierra de Líbar; N36°36′52′′ W05°23′16′′, 663 m; Quercus ilex, Ceratonia, 2007, StübenZFMK-DNA-JJ0261, ZFMK-TIS-cE0194berGU988244 GU987827
Onyxacalles bermejaensis Stüben 2001Spain: Málaga, NW Marbella, Sierra de las Nieves; N36°39′52′′ W05°04′57′′, 1043 m; Quercus ilex, Echinodera spinosa, 2009, StübenZFMK-DNA-JJ0747, ZFMK-TIS-cE778berGU988506 GU988066
Onyxacalles bermejaensis Stüben 2001Spain: Prov. Málaga, Algatocín, near Opayar; N36°34′39′′ W05°18′13′′, 576 m, Quercus sp., 17.8.2010, StübenZFMK-DNA-JJ1062, ZFMK-TIS-cES1062JN121398 _________
Onyxacalles gibraltarensis Stüben 2002Spain: Cádiz, 10 km SW Algeciras, El Bujeo; N36°04′10′′ W05°31′48′′, 257 m; Quercus suber, 2007, StübenZFMK-DNA-JJ0268, ZFMK-TIS-cE0206gibGU988249 GU987832
Onyxacalles gibraltarensis Stüben 2002Morocco: Rif, SW Oued-Laou, river, O. Laou; N35°17′47′′ W05°13′38′′, 210 m; Quercus suber, Smilax, Arbutus, 2009, StübenZFMK-DNA-JJ0854, ZFMK-TIS-cE897gibGU988577 GU988137
Onyxacalles gibraltarensis Stüben 2002Morocco: W Sebta, vir. Biutz; N35°53′04′′ W05°24′08′′, 337 m; Quercus suber, Smilax, Arbutus, 2009, StübenZFMK-DNA-JJ0856, ZFMK-TIS-cE899gibGU988578 GU988138
Onyxacalles gibraltarensis Stüben 2002Morocco: S Ksar-es-Seghir; N35°45′16′′ W05°30′49′′, 278 m; Pistacia, Quercus suber, 2009, StübenZFMK-DNA-JJ0864, ZFMK-TIS-cE907gibGU988584 GU988144
Onyxacalles gibraltarensis Stüben 2002Spain: Cádiz, Los Barrios, Alcornocales N.P., between Facinos, Río Las Cañas and Mantera Torero; Olea europaea, 2009, TorresZFMK-DNA-JJ0912, ZFMK-TIS-cE949gibGU988608 _________
Onyxacalles luigionii (A. & F. Solari 1907)Italy: Campania, Cilento, 6 km SE Vallo d. Lucania, M. Sacro o Gelbison; N40°12′41′′ E15°19′42′′, 1544 m; Fagus, 2008, StübenZFMK-DNA-JJ0561, ZFMK-TIS-cI622luiGU988407 GU987967
Onyxacalles luigionii (A. & F. Solari 1907)Italy: Campania, Monti Picentini, 9 km N Acerno, Piano Laceno; N40°48′58′′ E15°07′35′′, 1210 m; Fagus, 2008, StübenZFMK-DNA-JJ0574, ZFMK-TIS-cI635luiGU988417 GU987977
Onyxacalles luigionii (A. & F. Solari 1907)Italy: Basilicata, Monte Pollino, 9 km SE Rotonda, Rif. de Gasperi; N39°54′37′′ E16°07′15′′, 1486 m; Fagus, 2008, StübenZFMK-DNA-JJ0577, ZFMK-TIS-cI638luiGU988418 GU987979
Onyxacalles maginaensis Stüben 2004Spain: Andalucía, 28 km E Jaén, Sierra Magina; N37°43′21′′ W03°29′11′′, 1600 m; Quercus ilex, 2005, StübenZFMK-DNA-JJ0147, ZFMK-TIS-cE0169magEU286327 EU286491
Onyxacalles maginaensis Stüben 2004Spain: Almería, 11 km NW Laujar de Andarax, Sierra Nevada, Bayárcal; N37°02′27′′ W03°00′12′′, 1291 m; Quercus ilex, 2007, StübenZFMK-DNA-JJ0257, ZFMK-TIS-cE0187magsubmitted to GenBank
Onyxacalles maginaensis Stüben 2004Spain: Teruel, S. Javalambre, Fuente la Risca near Arcos de las Salinas; N39°59′56′′ W01°01′21′′, 1121 m; Amelanchier ovalis, Acer monspessulanum, Erinacea anthyllis, Ulex, 2008, StübenZFMK-DNA-JJ0541, ZFMK-TIS-cE602magGU988390 GU987950
Onyxacalles neglectus Kulbe 1999Spain: Canary Islands, La Gomera, S Hermigua, El Cedro, Las Mimbreras; N28°07′27′′ W17°13′26′′, 901 m; laurisilva, 2008, Astrin and StübenZFMK-DNA-JJ0640, ZFMK-TIS-cE713negFJ716525 GU988014
Onyxacalles nuraghi sp.n.Italy: W-Sardinia, E Macomer: above Lei; N40°19′54′′ E08°53′49′′, 1020 m; Quercus, Acer monspessulanum, 4.10.2010, StübenZFMK-DNA-JJ1090, ZFMK-TIS-cIT1090JN642097 JN121399
Onyxacalles nuraghi sp.n.Italy: W-Sardinia, E Macomer: above Lei; N40°19′17′′ E08°53′52′′, 586 m; Quercus ilex, 7.10.2010, StübenZFMK-DNA-JJ1097, ZFMK-TIS-cIT1097JN642098 JN121300
Onyxacalles portusveneris (Mayet 1903) [27]France: Gard, 15 km NE Nimes, Pont du Gard, Collias; N43°57′03′′ E04°28′59′′, 68 m; Quercus ilex, 2006, StübenZFMK-DNA-JJ0144, ZFMK-TIS-cF0166porEU286326 EU286490
Onyxacalles portusveneris (Mayet 1903) [27]Morocco: Rif Mts., 10 km W Ketama = Issague; N34°57′40′′ W04°40′51′′, 1600 m; Prunus lusitanica, 2001, StübenZFMK-DNA-JJ0375, ZFMK-TIS-cM480porGU988311 _________
Onyxacalles portusveneris (Mayet 1903) [27]Morocco: M-Atlas, 10 km S Ain-Leuh; N33°13′48′′ W05°20′50′′, 1700 m; Quercus ilex, Rubus, Cedrus, 2002, StübenZFMK-DNA-JJ0376, ZFMK-TIS-cM481porGU988312 _________
Onyxacalles portusveneris (Mayet 1903) [27]Morocco: High Atlas, E Marrakech, N Taddert, (near Tazouguerte); N31°28′07′′ W07°24′59′′, 1727 m; Quercus, 2009, StübenZFMK-DNA-JJ0846, ZFMK-TIS-cE889portGU988573 GU988133
Onyxacalles portusveneris (Mayet 1903) [27]Morocco: Middle Atlas, S Azrou, Äin Leuh; N33°16′50′′ W05°20′18′′, 1582 m; Quercus ilex, Euphorbia, 2009, StübenZFMK-DNA-JJ0847, ZFMK-TIS-cE890portGU988574 GU988134
Onyxacalles portusveneris (Mayet 1903) [27]Morocco: Middle Atlas, S Azrou, S Äin Leuh; N33°14′57′′ W05°21′04′′, 1715 m; Quercus ilex, 2009, StübenZFMK-DNA-JJ0848, ZFMK-TIS-cE891portGU988575 GU988135
Onyxacalles portusveneris (Mayet 1903) [27]Morocco: Rif, 10 km W Ketama; N34°57′40′′ W04°40′51′′, 1600 m; Cedrus, Prunus, 2009, StübenZFMK-DNA-JJ0869, ZFMK-TIS-cE912portGU988587 GU988147
Onyxacalles portusveneris (Mayet 1903) [27] (Onyxacalles sp.)Morocco: High Atlas, 59 km SE Marrakech; N31°28′19′′ W07°24′22′′, 1500 m; Quercus ilex, Quercus suber, 2002, StübenZFMK-DNA-JJ0472, ZFMK-TIS-cM482magGU988350 GU987922
Onyxacalles ringeli Kulbe 1999Spain: Canary Islands, La Palma, Cumbre Nueva, 4,5 km SE El Paso, El Pilar; N28°37′37′′ W17°49′45′′, 1432 m; laurisilva, 2006, StübenZFMK-DNA-JJ0087, ZFMK-TIS-cC0171rinEU286300 EU286465
Onyxacalles sp. 1 (O. luigionii)France: Alpes-Maritimes, 3 km W Sospel, Col de Braus; N43°52′34′′ E07°24′17′′, 1051 m; Quercus pubescens, Ostrya carpinifolia, broom, 2007, StübenZFMK-DNA-JJ0411, ZFMK-TIS-cF440luiGU988325 GU987897
Onyxacalles torre sp. n. (O. henoni)France: Corsica, Col de Vizzavona, 22 km S Corte; N42°06′45′′ E09°06′49′′, 1100 m; Fagus, 2001, StübenZFMK-DNA-JJ0882, ZFMK-TIS-cF479henGU988592 _________
Onyxacalles valencianus Germann 2005Spain: Alicante, 7 km SW Alcoi, Sierra de Menechaor, Santurio de la Font Roja; N38°39′34′′ W00°32′29′′,1296 m; Quercus ilex, 2007, StübenZFMK-DNA-JJ0172, ZFMK-TIS-cE0180valEU286331 EU286495
Onyxacalles valencianus Germann 2005Spain: Castellón, Morella, Barranco de la Bota; N40°33′12′′ W00°00′27′′, 814 m; Quercus ilex, Hedera helix, 2008, StübenZFMK-DNA-JJ0546, ZFMK-TIS-cE607valGU988393 GU987953
Onyxacalles valencianus Germann 2005Spain: Barcelona, above dry river bed, near Vallirana; N41°22′36′′ E01°55′02′′, 245 m; Quercus ilex, Ficus carica, Smilax aspera, 2008, StübenZFMK-DNA-JJ0551, ZFMK-TIS-cE612valGU988398 GU987958
Onyxacalles valencianus Germann 2005spain: Barcelona, S. Montseny, Tordera valley, near St. Marçal; N41°48′01′′ E02°25′15′′, 1060 m, 2008, StübenZFMK-DNA-JJ0553, ZFMK-TIS-cE614valGU988400 GU987960
Onyxacalles verrucosus (Wollaston 1863)Spain: Canary Islands, El Hierro, 7 km W La Frontera, Pista Derrabado; N27°44′29′′ W18°03′24′′, 895 m; Laurus azorica, 2006, StübenZFMK-DNA-JJ0137, ZFMK-TIS-cC0170verEU286324 EU286488
Onyxacalles verrucosus (Wollaston 1863)Spain: Canary Islands, Tenerife, 6 km N La Laguna, Monte de las Mercedes; N28°31′50′′ W16°17′09′′, 950 m; laurisilva, 2003, StübenZFMK-DNA-JJ0509, ZFMK-TIS-cE570ver_________ GU987937
Onyxacalles verrucosus (Wollaston 1863)Spain: Canary Islands, Tenerife, 4 km S Los Silos, Teno Mts., Monte del Aqua; N28°19′20′′ W16°49′14′′, 700 m; laurisilva, 2003, StübenZFMK-DNA-JJ0510, ZFMK-TIS-cE571verGU988373 GU987938
Onyxacalles verrucosus (Wollaston 1863)Spain: Canary Islands, Tenerife, NE La Laguna, Anaga Mts. near Moquinal; N28°31′55′′ W16°17′24′′, 840 m; laurisilva, 2008, Astrin and StübenZFMK-DNA-JJ0604, ZFMK-TIS-cE677verGU988433 GU987995
Onyxacalles verrucosus (Wollaston 1863)Spain: Canary Islands, Tenerife, NE La Laguna, Anaga Mts. near Chinobre; N28°33′21′′ W16°10′46′′, 808 m; Laurus, Ixanthus viscosus, 2008, Astrin and StübenZFMK-DNA-JJ0605, ZFMK-TIS-cE678verGU988434 GU987996
Onyxacalles verrucosus (Wollaston 1863)Spain: Canary Islands, Tenerife, SW Los Silos, Teno Mts., Monte del Agua, Chupadero; N28°19′23′′ W16°49′12′′, 940 m; Laurus novocanariensis, 2008, Astrin, Stüben, Behne and FlorenZFMK-DNA-JJ0776, ZFMK-TIS-cE813verGU988524 GU988085
Onyxacalles vilae sp.n. (O. luigionii)France: Isère, 2 km SE Lans en Vercors, Montagne de Lans; N45°06′45′′ E05°36′21′′, 1352 m; Abies, Fagus, Fraxinus, 2005, StübenZFMK-DNA-JJ0025, ZFMK-TIS-cI0027luiEU286286 EU286451
Onyxacalles vilae sp.n. (O. luigionii)Croatia: Dalmatian, 8 km E Karlobag, Velebit Mts., Stupacinovo; N44°32′41′′ E15°09′58′′, 1049 m; Fagus, 2007, StübenZFMK-DNA-JJ0195, ZFMK-TIS-cHR0339luiEU286348 EU286512