| | Cause of direct mortality | Geographic distribution of determinant | Locations where determinant reported | Selected references |
| | Hunting | Range-wide | Range-wide | [27, 28] | | Predation | Range-wide | Alberta, Canada, and Montana | [29, 30] | | Arizona | [31, 32] | | California | [33–36] | | Colorado | [27] | | Montana | [37] | | New Mexico | [38, 39] |
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