A Run-Length Encoding Approach for Path Analysis of C. elegans Search Behavior

Table 3

Run-length encoding descriptors.

Formula	Description
	Short run encoding (SRE) measures the distribution of short runs
	Long run encoding (LRE) measures the distribution of long runs
	Low angle run encoding (LARE) measures the distribution of the small angles
	High angle run encoding (HARE) measures the distribution of the large angles
	Short run low angle encoding (SRLAE) measures the joint distribution of short runs and sharp turns
	Short run high angle encoding (SRHAE) measures the joint distribution of short runs and shallow turns
	Large run low angle encoding (LRLAE) measures the joint distribution of large runs and sharp turns
	Large run low angle encoding (LRHAE) measures the joint distribution of large runs and shallow turns
	Angle level nonuniformity (ALN) measures the similarity of angle level distributions
	Run length nonuniformity (RLN) measures the similarity of run length distributions
	Run Percentage (RP) measures the homogeneity of the distribution of runs