Tree Diversity and Community Composition of the Tutong White Sands, Brunei Darussalam: A Rare Tropical Heath Forest Ecosystem

Table 1

Soil and environmental variables (mean ± SE) in six plots in the Tutong White Sands. and values from one-way ANOVA are shown for each variable. Different superscript letters within a column indicate significant differences at α = 0.05.

	Soil and environmental variables
Plot number	Total nitrogen (mg g^−1)	Total phosphorus (mg g^−1)	pH (units)	GWC (%)	Canopy openness (%)	Litter depth (m)
1	3.65 ± 2.20	0.12 ± 0.040	4.70 ± 0.27	47.3 ± 22.70	7.00 ± 1.15	0.52 ± 0.01
2	0.44 ± 0.23	0.04 ± 0.003	4.00 ± 0.04	66.8 ± 3.24	14.10 ± 1.26	0.39 ± 0.04
3	0.48 ± 0.32	0.03 ± 0.003	4.00 ± 0.11	73.1 ± 2.99	12.30 ± 3.03	0.33 ± 0.03
4	0.28 ± 0.06	0.03 ± 0.010	3.88 ± 0.11	88.7 ± 1.70	6.91 ± 1.37	0.15 ± 0.03
5	7.58 ± 1.20	0.14 ± 0.020	4.45 ± 0.03	19.6 ± 3.43	0.44 ± 0.17	0.57 ± 0.06
6	10.37 ± 0.44	0.17 ± 0.009	3.85 ± 0.28	14.7 ± 1.12	5.25 ± 0.82	0.56 ± 0.07
value	20.42	10.19	4.04	9.71	9.85	14.11
value	<0.001	<0.001	<0.05	<0.001	<0.001	<0.001