Modeling Aquatic Macroinvertebrate Richness Using Landscape Attributes

Table 4

Feeding guild, water quality tolerance, and life habit preferences for macroinvertebrate families in the study area and associated habitat classes.

Family	Feeding guild	Water quality tolerance	Habitat preferences	HCB	HCG	HOB	HOG	LCB	LCG	LOB	LOG
Aeolosomatidae	cf	T	bur	X	X	X	X	X	X	X	X
Ancylidae	sc	I	all		X		X
Asellidae	cg	T	all	X	X	X	X	X	X	X	X
Athericidae	pr	I	sp		X		X		X		X
Baetidae	cg	I	sw		X		X		X		X
Bithyniidae	sc	T	all	X	X	X	X
Blephariceridae	sc	I	clg		X		X
Brachycentridae	cf	I	clg		X		X
Caenidae	cg	I	sp		X		X		X		X
Calopterygidae	pr	I	clb		X		X		X		X
Cambaridae	cg	I	all		X		X		X		X
Capniidae	sh	I	sp		X				X
Carabidae	pr	I	clg		X		X
Ceratopogonidae	pr	I	sp		X		X		X		X
Chironomidae	cg	I	sp		X		X		X		X
Chloroperlidae	pr	I	clg		X		X
Coenagrionidae	pr	T	clb	X	X	X	X	X	X	X	X
Corixidae	pr	I	sw		X		X		X		X
Corydalidae	pr	I	clg		X		X
Curculionidae	sh	I	clg		X
Dixidae	cg	I	sw		X		X		X		X
Dolichopodidae	pr	I	sp		X		X		X		X
Dytiscidae	pr	I	clb		X		X		X		X
Elmidae	sc	I	clg		X		X
Empididae	pr	I	sp		X		X		X		X
Enchytraeidae	cg	T	bur	X	X	X	X	X	X	X	X
Ephemerellidae	cg	I	clg		X		X
Ephemeridae	cg	I	bur		X		X		X		X
Gammaridae	cg	I	all		X		X		X		X
Glossosomatidae	sc	I	clg		X		X
Gomphidae	pr	I	bur		X		X		X		X
Heptageniidae	sc	I	clg		X		X
Hydrobiidae	sc	T	all	X	X	X	X
Hydrophilidae	cg	I	sw		X		X		X		X
Hydropsychidae	cf	I	clg		X		X
Hydroptilidae	sc	I	clg		X		X
Lepidostomatidae	sh	I	clb		X				X
Leptoceridae	cg	I	clb		X		X		X		X
Leptophlebiidae	cg	I	sw		X		X		X		X
Leuctridae	sh	I	sp		X				X
Limnephilidae	sh	I	sp		X				X
Muscidae	pr	I	sp		X		X		X		X
Naididae	cg	T	bur	X	X	X	X	X	X	X	X
Nemouridae	sh	I	sp		X				X
Noteridae	pr	I	sw		X		X		X		X
Oligoneuriidae	cg	I	sw		X		X		X		X
Orthocladiinae	cg	T	bur	X	X	X	X	X	X	X	X
Peltoperlidae	sh	I	clg		X
Perlidae	pr	I	clg		X		X
Perlodidae	pr	I	clg		X		X
Philopotamidae	cf	I	clg		X		X
Phryganeidae	sh	I	clb		X				X
Physidae	cg	T	all	X	X	X	X	X	X	X	X
Pisidiidae	cf	I	bur		X		X		X		X
Polycentropodidae	cf	I	clg		X		X
Polymitarcyidae	cg	I	bur		X		X		X		X
Potamanthidae	cg	I	sp		X		X		X		X
Psephenidae	sc	I	clg		X		X
Psychomyiidae	cg	I	clg		X		X
Pteronarcyidae	sh	I	clg		X
Pyralidae	sh	I	clb		X				X
Rhyacophilidae	pr	I	clg		X		X
Saldidae	pr	I	clb		X		X		X		X
Sialidae	pr	I	bur		X		X		X		X
Simuliidae	cf	I	clg		X		X
Siphlonuridae	cg	I	sw		X		X		X		X
Sphaeriidae	cf	I	bur		X		X		X		X
Tabanidae	pr	I	sp		X		X		X		X
Talitridae	cg	T	bur	X	X	X	X	X	X	X	X
Tetrastemmatidae	pr	T	all	X	X	X	X	X	X	X	X
Tipulidae	sh	I	bur		X				X
Tricorythidae	cg	I	sp		X		X		X		X
Tubificidae	cg	T	bur	X	X	X	X	X	X	X	X

H = high gradient, L = low gradient, C = closed canopy, O = open canopy, G = water quality suitable for life support, and B = water quality biologically stressed;
cf = collector filterer, cg = collector gatherer, sc = scraper, sh = shredder, pr = predator; I = intolerant, T = tolerant; bur = burrower, clb = climber, clg = clinger, sp = sprawler, sw = swimmer, and all = all life habits represented.