Research Article

Common Expression Quantitative Trait Loci Shared by Histone Genes

Table 2

Associations of the identified eQTLs with expression of histone and nonhistone genes all over the human genome.

eQTLGeneGeneGeneGene

rs10919229HIST1H4B (6)2.51E−10HIST1H3J (6)3.40E−08HIST1H3H (6)2.02E−07SASH3 (X)2.89E−06
HIST1H2BM (6)4.38E−09HIST1H3B (6)4.64E−08HIST1H1E (6)2.50E−07HIST1H2AH (6)4.61E−06
HIST1H1B (6)1.81E−08RPPH1 (14)5.82E−08HIST1H2AB (6)2.54E−07
SCARNA13 (14)3.38E−08SCARNA12 (12)1.22E−07HIST1H2AM (6)9.76E−07

rs191508159SNORA53 (12)4.38E−13SCARNA2 (1)5.28E−08HIST2H3C (1)1.41E−07HIST1H3H (6)5.46E−07
HIST1H4A (6)3.41E−11HIST1H1E (6)7.02E−08HIST1H2AL (6)1.99E−07G3BP2 (4)5.51E−07
HIST1H2AJ (6)5.72E−09HIST2H2AB (1)7.35E−08HIST1H1B (6)2.67E−07HIST1H2AH (6)3.21E−06
HIST1H3J (6)3.33E−08HIST1H2AB (6)1.02E−07HIST1H4K (6)2.99E−07HIST1H3A (6)4.06E−06
HIST1H4J (6)4.63E−08HIST1H3B (6)1.14E−07HIST1H3I (6)5.12E−07HIST1H2BM (6)4.73E−06

rs13432631HIST1H4B (6)1.78E−11HIST1H3B (6)1.13E−08HIST1H2AJ (6)1.86E−07HIST1H2AH (6)6.84E−07
SCARNA12 (12)1.70E−10HIST1H2AB (6)2.83E−08HIST3H2BB (1)1.91E−07RPPH1 (14)7.69E−07
HIST1H2BM (6)1.95E−09HIST1H3I (6)1.01E−07HIST1H3H (6)2.56E−07HIST1H3A (6)2.10E−06
HIST1H1B (6)3.33E−09SCARNA13 (14)1.23E−07HIST1H4K (6)3.45E−07HIST1H4J (6)2.48E−06
HIST1H3J (6)1.11E−08HIST1H1E (6)1.74E−07HIST2H2AB (1)6.28E−07HIST1H4A (6)2.81E−06

rs189340111HIST1H4B (6)2.99E−12HIST1H3I (6)1.73E−09HIST1H4J (6)3.30E−08SCARNA2 (1)4.27E−07
HIST1H1B (6)2.91E−11HIST2H2AB (1)1.78E−09HIST1H3A (6)3.60E−08PIGF (2)6.15E−07
HIST1H1E (6)6.90E−11HIST1H2AJ (6)2.58E−09HIST1H4A (6)7.70E−08SCARNA13 (14)1.55E−06
HIST1H3J (6)8.17E−11HIST1H3H (6)3.93E−09HIST1H2AM (6)1.00E−07RPPH1 (14)1.91E−06
HIST1H2AH (6)2.34E−10HIST1H4K (6)9.71E−09SCARNA12 (12)1.05E−07HIST1H2AL (6)2.40E−06
HIST1H2BM (6)2.58E−10HIST1H2AB (6)1.81E−08SCARNA7 (3)1.81E−07
HIST1H3B (6)4.62E−10HIST3H2BB (1)3.14E−08HIST2H3C (1)2.90E−07

rs75372391HIST1H2BM (6)6.19E−12HIST1H3I (6)5.56E−09HIST1H4A (6)3.57E−08HIST1H2AG (6)1.77E−07
HIST1H4B (6)1.39E−11HIST1H2AJ (6)6.33E−09HIST1H2AH (6)5.13E−08HIST1H2AK (6)6.45E−07
HIST1H1B (6)2.92E−11HIST1H2AB (6)7.88E−09HIST1H2AM (6)5.45E−08SCARNA7 (3)1.95E−06
RPPH1 (14)1.53E−10HIST3H2BB (1)9.94E−09HIST1H4J (6)5.90E−08HIST1H3A (6)2.31E−06
HIST1H3B (6)2.00E−10HIST1H3H (6)1.10E−08HIST2H3C (1)9.35E−08SCARNA2 (1)2.85E−06
HIST1H1E (6)6.26E−10HIST2H2AB (1)2.33E−08SCARNA12 (12)9.42E−08
HIST1H3J (6)1.15E−09HIST1H4K (6)3.21E−08HIST1H2AL (6)1.09E−07

rs72853591HIST1H4B (6)9.68E−11SCARNA13 (14)1.03E−08SCARNA12 (12)2.12E−07HIST1H2AH (6)2.69E−06
HIST1H1B (6)5.88E−10HIST1H3J (6)1.17E−08HIST2H2AB (1)4.57E−07HIST1H2AL (6)2.72E−06
HIST1H2BM (6)6.02E−10HIST1H1E (6)1.88E−08HIST1H4A (6)1.16E−06HIST1H4J (6)3.52E−06
RPPH1 (14)1.96E−09HIST1H3H (6)1.56E−07HIST1H4K (6)1.57E−06HIST1H3A (6)4.33E−06
HIST1H3B (6)4.11E−09HIST1H2AJ (6)1.74E−07SNORD17 (20)1.66E−06HIST2H3C (1)4.67E−06
HIST1H3I (6)8.43E−09HIST1H2AB (6)1.84E−07SCARNA2 (1)2.24E−06

rs17801458RPPH1 (14)1.31E−15HIST1H2AL (6)9.35E−12SCARNA2 (1)9.35E−10HIST1H2AK (6)1.75E−06
HIST1H1B (6)1.62E−14HIST2H2AB (1)1.43E−11HIST1H3A (6)5.23E−09VGLL4 (3)1.88E−06
HIST1H2BM (6)3.12E−14HIST2H3C (1)4.10E−11HIST1H2AG (6)6.96E−09SNORA53 (12)2.09E−06
HIST1H3B (6)1.75E−13HIST1H4K (6)4.54E−11HIST1H2AH (6)1.34E−08WASF2 (1)3.49E−06
HIST1H4B (6)4.90E−13HIST1H4J (6)4.67E−11SNORD17 (20)2.36E−08UBE2R2 (9)4.47E−06
HIST1H2AB (6)8.94E−13HIST1H3H (6)5.05E−11SCARNA12 (12)2.60E−08
HIST1H3J (6)1.28E−12HIST1H3I (6)6.66E−11SCARNA7 (3)1.91E−07
HIST1H1E (6)3.33E−12HIST3H2BB (1)6.82E−11SCARNA13 (14)4.53E−07
HIST1H2AJ (6)4.01E−12HIST1H4A (6)1.04E−10HIST1H2AM (6)1.16E−06

rs11684178HIST1H2BM (6)6.71E−12HIST1H1E (6)3.59E−10HIST1H3A (6)2.78E−09NOL11 (17)4.61E−07
RPPH1 (14)7.92E−12HIST1H4J (6)3.64E−10HIST2H3C (1)3.55E−09SCARNA12 (12)8.16E−07
HIST1H1B (6)2.56E−11HIST1H4B (6)3.69E−10HIST3H2BB (1)9.84E−09SCARNA13 (14)1.09E−06
HIST1H3B (6)6.72E−11HIST1H2AB (6)4.40E−10HIST1H4A (6)2.03E−08DDX17 (22)1.32E−06
HIST2H2AB (1)1.04E−10HIST1H2AH (6)7.01E−10HIST1H4K (6)3.26E−08MRPS24 (7)3.22E−06
HIST1H3J (6)1.10E−10HIST1H2AJ (6)8.58E−10SCARNA7 (3)3.37E−08FASTKD2 (2)3.42E−06
HIST1H2AL (6)1.67E−10HIST1H3H (6)1.56E−09HIST1H2AG (6)3.80E−08SNORD17 (20)3.99E−06
HIST1H3I (6)3.41E−10SCARNA2 (1)2.47E−09HIST1H2AK (6)7.75E−08EFTUD2 (17)4.04E−06

rs6718590HIST1H2BM (6)1.15E−10HIST1H1E (6)2.68E−09HIST1H3H (6)1.57E−08NOL11 (17)4.12E−07
RPPH1 (14)1.92E−10HIST1H3I (6)3.57E−09HIST1H3A (6)1.98E−08SCARNA12 (12)1.10E−06
HIST1H1B (6)3.63E−10HIST1H4B (6)4.90E−09SCARNA2 (1)2.30E−08APPBP2 (17)1.77E−06
HIST1H2AL (6)4.03E−10HIST1H2AB (6)6.23E−09HIST1H2AG (6)8.36E−08ATP5A1 (18)3.68E−06
HIST2H2AB (1)5.83E−10HIST1H2AH (6)6.89E−09HIST1H4A (6)1.92E−07PITPNB (22)3.92E−06
HIST1H3J (6)9.78E−10HIST1H4J (6)9.48E−09SCARNA7 (3)2.51E−07
HIST3H2BB (1)1.30E−09HIST2H3C (1)1.01E−08HIST1H4K (6)3.19E−07
HIST1H3B (6)1.74E−09HIST1H2AJ (6)1.10E−08HIST1H2AK (6)3.22E−07

rs111875224HIST1H2AM (6)1.24E−11SCARNA7 (3)2.79E−08HIST1H3H (6)2.63E−07GTF2E2 (8)2.28E−06
HIST1H4B (6)8.51E−11HIST3H2BB (1)3.71E−08SCARNA2 (1)3.89E−07C18orf32 (18)2.68E−06
HIST1H1B (6)7.63E−10HIST1H3J (6)9.07E−08HIST1H2AJ (6)4.51E−07CD63 (12)2.80E−06
HIST1H1E (6)1.11E−09SCARNA12 (12)1.02E−07SCARNA13 (14)4.99E−07SERINC1 (6)3.24E−06
HIST1H2BM (6)1.41E−09HIST1H3I (6)1.14E−07HIST1H3A (6)9.59E−07
HIST1H3B (6)1.41E−08HIST1H2AB (6)1.32E−07HIST1H2AL (6)1.60E−06
HIST1H2AH (6)2.27E−08HIST2H2AB (1)1.38E−07RPPH1 (14)1.87E−06

rs849577HIST1H4A (6)4.48E−11HIST1H1E (6)3.42E−08HIST1H3I (6)4.33E−07HIST1H3A (6)2.03E−06
SNORA53 (12)3.16E−10HIST2H2AB (1)4.00E−08HIST1H1B (6)5.65E−07HIST1H3H (6)2.06E−06
HIST1H4J (6)8.79E−10HIST1H3B (6)8.53E−08HIST1H3J (6)6.70E−07RPPH1 (14)3.22E−06
SCARNA2 (1)1.23E−09HIST1H2AL (6)1.35E−07HIST1H2AH (6)6.75E−07HIST1H2BM (6)4.42E−06
HIST1H4K (6)2.46E−09HIST2H3C (1)2.09E−07SCARNA7 (3)1.71E−06
HIST1H2AJ (6)1.12E−08HIST1H2AB (6)3.52E−07SNORD17 (20)1.77E−06

rs830612HIST1H4B (6)1.28E−10HIST1H3B (6)1.16E−08SNORD17 (20)1.52E−07RPPH1 (14)6.10E−07
HIST1H1B (6)6.19E−10HIST1H3H (6)1.67E−08SASH3 (X)1.74E−07HIST1H2AJ (6)6.67E−07
HIST1H2BM (6)2.66E−09HIST1H1E (6)2.00E−08HIST1H2AH (6)2.54E−07SCARNA7 (3)1.56E−06
HIST1H3J (6)1.05E−08HIST1H3I (6)4.17E−08HIST1H3A (6)3.80E−07HIST1H2AB (6)3.15E−06

rs150958961HIST1H4A (6)8.77E−11HIST1H1B (6)4.03E−09HIST1H4K (6)3.84E−08HIST2H2AB (1)4.70E−07
HIST1H3J (6)3.41E−10SCARNA2 (1)6.96E−09HIST1H3H (6)4.81E−08SCARNA7 (3)5.49E−07
HIST1H2AJ (6)3.78E−10HIST1H2AH (6)1.72E−08HIST1H3I (6)1.01E−07HIST1H3A (6)1.04E−06
HIST1H2AB (6)3.81E−10HIST1H2AL (6)1.86E−08HIST1H4B (6)1.38E−07SNORA57 (11)1.66E−06
HIST1H3B (6)4.54E−10HIST1H2BM (6)2.30E−08HIST3H2BB (1)1.62E−07HIST1H2AG (6)2.46E−06
RPPH1 (14)7.11E−10HIST1H4J (6)2.31E−08SNORA53 (12)4.22E−07
HIST1H1E (6)1.80E−09HIST2H3C (1)2.57E−08SNORD17 (20)4.60E−07

rs148619316HIST1H4B (6)4.81E−11HIST1H3H (6)6.81E−08HIST1H2AJ (6)1.23E−06PPM1G (2)2.97E−06
RPPH1 (14)3.50E−09HIST1H1E (6)1.11E−07HIST1H3I (6)1.33E−06THRAP3 (1)3.14E−06
HIST1H1B (6)4.92E−09HIST1H2AH (6)1.66E−07SNORD17 (20)1.52E−06SUPT5H (19)3.91E−06
HIST1H3J (6)5.46E−09DDX28 (16)5.22E−07SCARNA12 (12)1.68E−06ZNF207 (17)4.48E−06
HIST1H3B (6)9.95E−09HIST1H2AB (6)6.80E−07HIST1H2AK (6)1.69E−06
HIST1H2BM (6)1.70E−08HIST2H2AB (1)1.14E−06C20orf11 (20)2.61E−06
SASH3 (X)3.00E−08HIST3H2BB (1)1.22E−06SCARNA7 (3)2.67E−06

rs6861945HIST1H3J (6)2.75E−10HIST1H4J (6)1.67E−08HIST1H3I (6)7.68E−08HIST1H2AK (6)1.18E−06
HIST1H3B (6)4.30E−09HIST1H1E (6)2.30E−08HIST1H2AB (6)1.38E−07HIST2H2AB (1)1.32E−06
HIST1H4B (6)4.44E−09HIST1H4K (6)3.40E−08SCARNA2 (1)1.55E−07SCARNA7 (3)2.25E−06
HIST1H2BM (6)7.98E−09HIST1H2AH (6)3.78E−08HIST1H4A (6)1.55E−07SNORD17 (20)2.57E−06
HIST1H1B (6)1.04E−08HIST1H3H (6)4.52E−08SASH3 (X)4.33E−07HIST1H2AL (6)4.34E−06
RPPH1 (14)1.11E−08HIST1H2AJ (6)5.09E−08HIST1H3A (6)8.63E−07

rs61564830HIST1H3J (6)2.62E−10SCARNA2 (1)5.85E−09HIST1H3I (6)3.37E−08HIST1H2AB (6)2.54E−06
HIST1H2AH (6)8.25E−10HIST1H1B (6)6.35E−09HIST1H2BM (6)3.45E−08HIST1H2AL (6)2.85E−06
HIST1H1E (6)9.74E−10HIST1H2AJ (6)1.00E−08SCARNA13 (14)1.68E−07UBE2R2 (9)4.53E−06
HIST1H3B (6)1.50E−09SNORD17 (20)1.23E−08RPPH1 (14)3.47E−07
HIST1H3H (6)1.58E−09SCARNA7 (3)2.46E−08HIST2H2AB (1)2.23E−06
HIST1H4B (6)2.05E−09HIST1H4A (6)2.75E−08MRPS31 (13)2.36E−06

rs144499291HIST1H4A (6)2.88E−10HIST1H2AK (6)4.16E−09HIST1H4K (6)9.65E−08FAM122A (9)2.05E−06
HIST1H1B (6)4.57E−10HIST1H3H (6)5.46E−09KPNB1 (17)1.48E−07RPPH1 (14)3.45E−06
HIST1H2AJ (6)7.99E−10HIST1H3I (6)1.06E−08SNORD17 (20)1.49E−07HIST3H2BB (1)4.06E−06
HIST1H1E (6)9.23E−10HIST1H2AB (6)1.12E−08HIST2H3C (1)2.10E−07
HIST1H2BM (6)1.15E−09HIST2H2AB (1)3.99E−08HIST1H4J (6)2.73E−07
HIST1H3J (6)2.45E−09HIST1H4B (6)4.58E−08SCARNA7 (3)2.83E−07
HIST1H3B (6)2.85E−09SCARNA2 (1)5.61E−08HIST1H2AL (6)4.25E−07
HIST1H2AH (6)4.05E−09SNORA53 (12)9.55E−08HIST1H3A (6)5.03E−07

rs71560740HIST1H2AJ (6)1.48E−10HIST1H3I (6)6.75E−09RPPH1 (14)1.71E−08HIST1H2AH (6)4.57E−07
HIST1H3A (6)2.41E−10HIST2H2AB (1)9.23E−09SNORA53 (12)2.32E−08HIST1H3H (6)9.55E−07
HIST1H2AB (6)8.54E−10HIST1H4J (6)9.80E−09HIST1H3J (6)2.54E−08TDRD12 (19)3.26E−06
HIST2H3C (1)1.32E−09HIST1H2BM (6)1.13E−08HIST1H1E (6)3.18E−08HIST1H2AM (6)3.74E−06
HIST1H3B (6)3.46E−09HIST1H4A (6)1.27E−08HIST1H2AL (6)7.54E−08HIST1H4B (6)3.78E−06
HIST1H1B (6)6.55E−09HIST1H4K (6)1.29E−08SCARNA2 (1)2.35E−07

rs118052517HIST1H3J (6)8.16E−13HIST1H4K (6)2.14E−10HIST1H4B (6)2.72E−09HIST3H2BB (1)3.49E−08
HIST1H4A (6)1.89E−12SCARNA2 (1)2.30E−10HIST1H2AH (6)2.81E−09LOXL4 (10)1.35E−06
HIST1H2BM (6)2.40E−12HIST1H4J (6)2.62E−10HIST2H3C (1)6.12E−09KRT8 (12)1.53E−06
HIST1H2AJ (6)3.64E−12HIST1H3I (6)2.77E−10SNORA53 (12)8.62E−09HIST1H2AM (6)1.63E−06
HIST1H3B (6)6.16E−12HIST1H3H (6)2.95E−10HIST1H3A (6)1.04E−08TRNP1 (1)1.82E−06
HIST1H1B (6)1.39E−11RPPH1 (14)1.53E−09HIST1H2AK (6)1.27E−08SCUBE2 (11)4.20E−06
HIST1H1E (6)1.78E−11HIST1H2AL (6)1.73E−09SNORD17 (20)1.61E−08
HIST1H2AB (6)1.16E−10HIST2H2AB (1)2.34E−09SCARNA7 (3)3.31E−08

rs77957383HIST1H4B (6)1.85E−10HIST3H2BB (1)4.38E−09SCARNA12 (12)1.82E−07CDC5L (6)3.27E−06
RPPH1 (14)3.19E−10HIST1H1E (6)2.17E−08SCARNA2 (1)2.84E−07HIST2H3C (1)4.43E−06
HIST1H2BM (6)4.46E−10HIST2H2AB (1)2.20E−08HIST1H3H (6)7.97E−07
HIST1H3B (6)1.04E−09HIST1H2AJ (6)2.95E−08HIST1H2AK (6)1.03E−06
HIST1H1B (6)1.38E−09SCARNA7 (3)3.95E−08HIST1H4A (6)1.61E−06
HIST1H3J (6)3.57E−09HIST1H3I (6)4.73E−08SCARNA13 (14)2.67E−06
HIST1H2AB (6)3.98E−09HIST1H2AH (6)1.49E−07HIST1H2AL (6)3.22E−06

rs113936625HIST1H4B (6)2.76E−10SCARNA13 (14)1.12E−07HIST1H3H (6)3.64E−07RPPH1 (14)2.97E−06
HIST1H2BM (6)2.29E−09SCARNA12 (12)1.20E−07HIST1H2AJ (6)9.36E−07HIST1H2AL (6)3.00E−06
HIST1H1B (6)8.82E−09HIST1H3B (6)1.36E−07HIST2H2AB (1)1.25E−06HIST1H3A (6)3.44E−06
HIST1H2AM (6)1.42E−08HIST1H2AB (6)1.79E−07HIST1H3I (6)1.41E−06SNORD17 (20)3.63E−06
HIST1H3J (6)3.32E−08HIST1H1E (6)2.11E−07HIST1H2AH (6)2.45E−06

rs75201173HIST1H2BM (6)1.45E−11HIST1H2AK (6)2.05E−09STYX (14)4.33E−07HMHA1 (19)2.49E−06
RPPH1 (14)6.79E−11HIST1H2AB (6)3.62E−09PRKAR1A (17)4.54E−07OSBPL11 (3)2.57E−06
HIST2H3C (1)1.22E−10SCARNA2 (1)1.15E−08SMAD2 (18)5.85E−07FBXL12 (19)2.73E−06
HIST1H1B (6)1.75E−10HIST1H2AL (6)1.56E−08EFTUD2 (17)6.59E−07FNBP1 (9)2.92E−06
HIST1H3B (6)1.94E−10HIST2H2AB (1)3.33E−08ZFP91 (11)8.19E−07SLC44A2 (19)3.26E−06
HIST1H4B (6)3.04E−10HIST1H3I (6)5.13E−08GBF1 (10)1.03E−06SCARNA12 (12)3.62E−06
HIST1H3H (6)6.76E−10HIST1H2AH (6)5.81E−08APBA3 (19)1.15E−06SCARNA13 (14)4.24E−06
HIST1H2AJ (6)6.95E−10HIST1H2AG (6)7.13E−08COX15 (10)1.90E−06ACLY (17)4.49E−06
SCARNA7 (3)1.45E−09HIST1H4K (6)1.46E−07SERBP1 (1)2.00E−06
HIST1H2AM (6)1.61E−09HIST3H2BB (1)1.64E−07HIST1H2BN (6)2.09E−06
HIST1H1E (6)1.79E−09HIST1H4A (6)3.81E−07HIST1H3A (6)2.19E−06
HIST1H3J (6)1.92E−09HIST1H4J (6)4.30E−07HIST1H2BO (6)2.22E−06

rs113659777HIST1H2BM (6)2.89E−11HIST1H1E (6)1.72E−09SNORD17 (20)1.14E−07SCARNA12 (12)1.62E−06
RPPH1 (14)7.27E−11HIST1H4K (6)3.31E−09HIST3H2BB (1)1.27E−07HTATSF1 (X)2.97E−06
HIST1H4B (6)8.17E−11HIST1H2AJ (6)5.31E−09HIST1H2AK (6)1.40E−07AGGF1 (5)3.52E−06
HIST1H1B (6)3.27E−10HIST1H2AB (6)7.80E−09SCARNA13 (14)2.84E−07MTFMT (15)3.89E−06
HIST1H3B (6)6.97E−10HIST1H2AL (6)1.05E−08HIST1H4A (6)3.39E−07
HIST2H2AB (1)9.75E−10SCARNA2 (1)1.72E−08HIST1H3I (6)3.63E−07
HIST1H2AM (6)9.86E−10HIST1H4J (6)4.06E−08HIST1H2AG (6)4.70E−07
HIST1H3J (6)1.08E−09HIST1H2AH (6)9.03E−08HIST1H3A (6)4.83E−07
HIST1H3H (6)1.37E−09HIST2H3C (1)9.54E−08SCARNA7 (3)1.13E−06

rs184239416HIST1H2BM (6)1.56E−12HIST2H2AB (1)2.30E−09SCARNA12 (12)9.43E−08HIST2H3C (1)1.42E−06
RPPH1 (14)2.76E−12HIST1H3H (6)2.98E−09HIST1H4A (6)9.92E−08GNAI3 (1)1.60E−06
HIST1H4B (6)9.13E−12HIST1H2AL (6)3.09E−09SCARNA13 (14)1.11E−07MAPRE1 (20)1.73E−06
HIST1H1B (6)2.84E−11HIST1H2AJ (6)3.83E−09HIST1H3I (6)1.46E−07RIOK3 (18)2.38E−06
HIST1H3J (6)3.88E−11HIST1H2AM (6)8.62E−09CAPZA1 (1)1.84E−07HIST1H2AK (6)2.55E−06
HIST1H3B (6)1.17E−10HIST1H2AB (6)1.50E−08HIST1H2AG (6)2.41E−07AGGF1 (5)2.75E−06
HIST1H1E (6)2.06E−10SCARNA2 (1)3.83E−08HIST1H3A (6)2.62E−07ARL6IP6 (2)3.00E−06
HIST3H2BB (1)2.37E−10HIST1H2AH (6)6.24E−08HIST1H4K (6)4.45E−07
SNORD17 (20)1.19E−09SCARNA7 (3)6.81E−08HIST1H4J (6)4.76E−07

rs41392245HIST1H2BM (6)4.19E−11HIST1H1E (6)3.18E−09HIST1H4K (6)2.14E−07HMHA1 (19)2.79E−06
RPPH1 (14)1.07E−10HIST1H2AB (6)4.10E−09PRKAR1A (17)3.46E−07FBXL12 (19)3.05E−06
HIST2H3C (1)1.34E−10HIST1H2AK (6)1.08E−08ZFP91 (11)3.49E−07SNORD17 (20)3.06E−06
HIST1H3B (6)2.23E−10SCARNA2 (1)1.47E−08SCARNA12 (12)4.21E−07APBA3 (19)3.07E−06
HIST1H1B (6)2.90E−10HIST1H2AL (6)1.89E−08STYX (14)7.62E−07HIST1H2BN (6)3.26E−06
HIST1H4B (6)3.59E−10HIST2H2AB (1)2.27E−08HIST1H4J (6)7.71E−07RDH5 (12)3.79E−06
HIST1H3H (6)1.08E−09HIST1H2AG (6)2.99E−08HIST1H4A (6)9.00E−07ACLY (17)4.07E−06
HIST1H2AJ (6)1.10E−09HIST1H2AH (6)4.46E−08HIST1H2BO (6)9.37E−07ZNF622 (5)4.27E−06
SCARNA7 (3)1.16E−09HIST1H3I (6)7.59E−08SMAD2 (18)1.58E−06
HIST1H2AM (6)2.39E−09HIST3H2BB (1)1.42E−07HIST1H3A (6)1.82E−06
HIST1H3J (6)2.76E−09COX15 (10)1.96E−07EFTUD2 (17)2.57E−06

rs11033879SNORA53 (12)2.29E−13HIST1H3J (6)2.08E−11HIST1H3A (6)4.97E−10HIST1H2AG (6)2.23E−07
HIST1H2AJ (6)2.53E−13HIST1H4K (6)2.16E−11RPPH1 (14)1.18E−09OBFC2B (12)1.22E−06
HIST1H1E (6)9.71E−13HIST1H2AB (6)3.14E−11HIST3H2BB (1)1.51E−09RUNDC1 (17)1.82E−06
HIST1H3B (6)9.80E−13HIST1H4J (6)4.85E−11HIST1H4B (6)2.07E−09NHS (X)1.92E−06
HIST2H2AB (1)1.12E−12HIST1H3I (6)7.46E−11HIST1H2AK (6)6.39E−09SVIL (10)4.02E−06
HIST1H1B (6)1.18E−12SCARNA2 (1)1.10E−10HIST1H2AL (6)1.64E−08
HIST1H4A (6)1.62E−12HIST1H3H (6)1.28E−10SCARNA7 (3)3.12E−08
HIST1H2BM (6)4.53E−12HIST1H2AH (6)2.70E−10FN1 (2)3.67E−08
HIST2H3C (1)1.15E−11SNORD17 (20)4.02E−10HIST1H2AM (6)1.68E−07

rs80143982HIST1H4A (6)2.65E−10NOTCH2NL (1)2.32E−07HIST1H3I (6)1.73E−06ZNF207 (17)4.57E−06
HIST3H2BB (1)3.31E−10RNF26 (11)3.73E−07HIST1H2AJ (6)2.16E−06
SNORA53 (12)2.42E−08SCARNA2 (1)6.10E−07SNORD17 (20)2.25E−06
HIST1H1E (6)3.15E−08HIST2H3C (1)1.51E−06MRFAP1L1 (4)4.02E−06

rs75349221HIST1H4A (6)2.75E−10HIST1H2BM (6)8.36E−09HIST1H3H (6)1.11E−07HIST1H2AB (6)1.91E−06
HIST1H1E (6)4.32E−10HIST1H3I (6)8.42E−09SCARNA2 (1)1.45E−07NDUFA8 (9)2.23E−06
HIST1H1B (6)7.02E−10SNORD17 (20)1.32E−08HIST1H4K (6)1.59E−07NDUFS1 (2)2.90E−06
HIST1H3B (6)1.26E−09SNORA53 (12)1.96E−08HIST1H4J (6)2.64E−07RPPH1 (14)2.95E−06
HIST1H2AJ (6)4.14E−09HIST1H4B (6)2.78E−08UBAP1 (9)3.35E−07TEX261 (2)3.35E−06
SCARNA7 (3)7.10E−09HIST2H2AB (1)5.63E−08HIST2H3C (1)1.36E−06RUNDC1 (17)3.63E−06
HIST1H3J (6)8.33E−09HIST1H2AH (6)8.54E−08SCARNA13 (14)1.61E−06HIST1H2AL (6)4.38E−06

rs72846811HIST1H4A (6)1.90E−11HIST1H3I (6)1.06E−07SCARNA7 (3)3.49E−07HIST1H3A (6)2.08E−06
SNORA53 (12)1.90E−10SCARNA2 (1)1.22E−07HIST1H3J (6)5.24E−07HIST1H2BM (6)2.77E−06
HIST1H1E (6)8.56E−09HIST1H2AH (6)2.32E−07HIST2H3C (1)1.12E−06HIST2H2AB (1)2.79E−06
HIST1H2AJ (6)7.92E−08HIST1H3B (6)2.44E−07HIST1H1B (6)1.24E−06

rs183898825SNORA53 (12)9.65E−11HIST1H4J (6)2.52E−08HIST1H1E (6)2.74E−07HIST1H2AH (6)7.54E−07
HIST1H4A (6)2.60E−10HIST1H2AJ (6)4.29E−08HIST2H2AB (1)4.28E−07SNORA70 (X)8.38E−07
HIST1H4K (6)7.93E−09HIST2H3C (1)8.18E−08HIST1H3B (6)4.88E−07HIST1H1B (6)1.58E−06
HIST1H3I (6)1.64E−08SCARNA7 (3)1.21E−07HIST1H3H (6)4.96E−07HIST1H2AB (6)2.66E−06
SCARNA2 (1)1.69E−08SNORD17 (20)1.78E−07HIST1H3A (6)5.65E−07HIST1H3J (6)3.18E−06

rs57185274HIST1H4B (6)2.04E−10HIST1H3J (6)4.54E−08HIST1H2AB (6)5.47E−07EPCAM (2)2.65E−06
HIST1H2BM (6)3.32E−09HIST1H3B (6)7.51E−08HIST1H3H (6)6.20E−07HIST2H3C (1)2.79E−06
HIST1H2AM (6)5.05E−09SCARNA12 (12)2.88E−07HIST1H2AJ (6)7.13E−07FNBP1 (9)3.03E−06
HIST1H1B (6)7.66E−09SCARNA13 (14)4.61E−07HIST2H2AB (1)9.68E−07HIST3H2BB (1)3.24E−06
HIST1H1E (6)2.02E−08HIST1H3I (6)5.12E−07HIST1H2AH (6)1.69E−06ZNF524 (19)4.57E−06

Figure in parentheses indicates chromosome number.