Spatial Dispersal of Douglas-Fir Beetle Populations in Colorado and Wyoming

Table 1

Total infestation areas for all years and regions shown with values of standard dispersal distance (SDD). Regional averages are displayed at the bottom.

Year	Colorado; eastern slope		Southwestern Colorado		Northwestern Colorado		Wyoming
	Area (ha)	SDD (m)	Area (ha)	SDD (m)	Area (ha)	SDD (m)	Area (ha)	SDD (m)
1994	7,135	903	702	1,996	0	—	0	—
1995	416	3,601	325	4,499	15	—	1,717	3,097
1996	290	1,037	2,227	1,288	5,607	—	490	656
1997	528	1,102	1,004	381	3	—	1,052	525
1998	3,843	2,718	1,218	—	255	204	214	667
1999	1,848	1,993	482	509	57	258	1,912	410
2000	2,174	1,680	3,030	792	172	342	2,977	1,542
2001	2,605	2,717	809	736	340	1,521	4,765	2,863
2002	2,350	—	3,312	1,770	175	535	10,970	894
2003	6,302	1,897	11,728	873	1,501	644	7,242	2,287
2004	7,304	1,641	8,869	2,486	1,049	2,532	27,571	3,739
2005	3,214	3,693	7,551	672	1,274	—	67,744	10,605
2006	4,644	970	5,630	2,389	3,233	1,384	6,754	1,008
2007	2,348	1,281	12,678	1,949	4,664	1,413	24,053	871
2008	555	3,530	8,122	804	2,711	315	8,535	3,817
2009	2,187	1,304	6,317	762	760	564	1,448	7,599
2010	5,108	2,124	8,983	708	757	601	2,938	2,030
Avg	3,109	2,012	4,882	1,390	1,328	859	10,022	2,663