Genome Sizes in Hepatica Mill: (Ranunculaceae) Show a Loss of DNA, Not a Gain, in Polyploids

Table 1

Hepatica species with their nuclear DNA content in pg, average for the species, standard deviation, and origin.

Hepatica species	2C DNA	Average	Standard	Origin
	in pg		deviation	(as = received as)
Section Hepatica
H. nobilis Schreber	32.7	33.0	0.9	Sweden
H. nobilis Schreber	32.7			Poland
H. nobilis Schreber	33.2			hort. blue flower
H. nobilis Schreber	32.3			hort. pink flower
H. nobilis Schreber	32.9			hort. white flower
H. nobilis Schreber	32.7			hort. Rubra plena
H. nobilis Schreber	33.7			Sweden, forma Plena
H. nobilis Schreber	33.3			“Crenatiloba”
H. nobilis Schreber	33.9			Marmor Crenata'
H. nobilis Schreber	33.3			Walter Otto' (double)
H. nobilis Schreber	33.2			as var glabrata, Sweden
H. nobilis Schreber	33.2			as var glabrata, S.Oeland
H. nobilis Schreber	32.3			as var pyrenaica
H. americana ssp. acuta (Pursh) Zonn.	34.6	34.8	1.0	USA, on limestone
H. americana ssp. americana (DC.) Ker Gawl.	34.9			USA, on acid soil
H. maxima (Nakai) Nakai	35.9	34.8	2.2	Korea, BSWJ 4344
H. maxima (Nakai) Nakai	34.0			Korea
H. maxima (Nakai) Nakai	34.4			Korea, Ullung Do
H. asiatica Nakai ssp. asiatica	36.6	36.3	1.4	China
H. asiatica Nakai ssp. asiatica	35.9			Korea
H. asiatica ssp. insularis (Nakai) Zonn.	34.7	35.4	1.5	Korea, BSWJ 859
H. asiatica ssp. insularis (Nakai) Zonn.	36.1			Korea, Cheju Do
H. asiatica ssp. japonica (Nakai) Zonn.	35.6	35.8	0.6	Japan, blue flower
H. asiatica ssp. japonica (Nakai) Zonn.	36.4			Japan, pink flower
H. asiatica ssp. japonica (Nakai) Zonn.	35.4			Japan
H. asiatica ssp. pubescens (Hiroe) Zonn.	70.0	70.0	2.9	Japan
H. asiatica ssp. pubescens (Hiroe) Zonn.	79.9	79.9	0.4	Japan, “Tenjinbai”
Section Angulosa (Ulbr.) Nakai
H. falconeri (Thomson) Steward	25.9*			Kashmir, Pakistan
H. henryi Steward	53.2	53.0	2.9	China
H. henryi Steward	52.7			via E. Myrholt
H. transsilvanica Fuss	55.5	54.2	2.5	Lilacina'
H. transsilvanica Fuss	54.4			Bulgaria
H. transsilvanica Fuss	54.2			Elison Spence'
H. transsilvanica Fuss	54.2			Loddon Blue'
H. transsilvanica Fuss	52.9
H. yamatutae Nakai	58.2	58.3	2.1	China, Emei Shan
H. yamatutae Nakai	59.5			China, “Marmorata”
H. yamatutae Nakai	58.9			China, black leaf
H. yamatutae Nakai	56.4			China, Emei Shan

*Recalculated from [7]