Research Article

Identification of Fine-Scale Marine Benthic Ecoclines by Multiple Parallel Ordination

Table 1

Sampling of sites based on stratified random sampling of geomorphological features. The classification into geomorphological features was based on a slightly modified version of an algorithm by Lundblad et al. [23], which takes into account fine- and broad-scaled geomorphological variation.

GeomorphologyCover (%)SitesBPI65BPI33Slope65Slope33Depth

Fine-scaled features
 Narrow depression 6.41-2, 26≤−1 SD≤–1 SD
 Broad depression with open bottom 3.23-4≤−1 SDw/in 1 SD
 Narrow crest7.65-6, 25≥1 SD≥1 SD
 Local depression on flat0.17w/in 1 SD≤–1 SD≤5
 Lateral midslope depression1.48-9w/in 1 SD≤–1 SD>5
 Local crest on flat0.110w/in 1 SD≥1 SD≤5
 Lateral midslope crest 1.211-12w/in 1 SD≥1 SD>5
 Shallow flat 28.413–15, 27>−1 SDw/in 1 SD≤5≤58.6
 Deep flat 15.716–18>−1 SDw/in 1 SD≤5>58.6
 Shallow slope 21.919–21, 28>−1 SDw/in 1 SD>5 and ≤70≤58.6
 Deep slope 14.022–24>−1 SDw/in 1 SD>5 and ≤70>58.6
 Total100.0
Broad-scaled features
 Depression 9.61–4, 26≤−1 SD
 Crest 11.15-6, 25≥1 SD
 Deep flat 16.17, 11, 16–18 w/in 1 SD≤5>58.6
 Shallow slope 19.68-9, 19–21, 28w/in 1 SD>5≤58.6
 Shallow flat 29.410, 13–15, 27w/in 1 SD≤5≤58.6
 Deep slope 14.212, 22–24 w/in 1 SD>5>58.6
 Total 100.0

Cover: estimated percentage of the study area covered by the feature; Sites: number of the sampled sites belonging to each fine- and broad-scaled feature class; BPI65 and BPI33: bathymetric position index rasters calculated from 65 and 33 m moving windows; Slope65 and Slope33: corresponding rasters for slope. The mean depth of the study, 58.6 m, was used to differentiate between shallow or deep slopes and flats, respectively.