
Variable  Definition 

 Number of susceptible female deer at time 
 Number of susceptible male deer at time 
 Number of exposed female deer at time 
 Number of exposed male deer at time 
 Number of infected female deer at time 
 Number of infected male deer at time 
 Fecundity 
 Transmission rate 
 Harvesting rate of females 
 Harvesting rate of males 
 Transmissibility to females from an asymptomatic deer 
 Transmissibility to females from infected deer 
 Transmissibility to males from asymptomatic deer 
 Transmissibility to males from infected deer 
 Rate of transmission from asymptomatic deer 
 Rate of transmission from infected deer 
 Rate of contact between infected males and susceptible females 
 Rate of contact between infected females and susceptible males 
 Rate of contact between infected males and susceptible males 
 Rate of contact between infected females and susceptible females 
 Incubation period 
 Lifespan for exposed deer 
 Lifespan for asymptomatic deer 
 Rate of progression from exposed class to infected class 
 Rate of progression from exposed class to asymptomatic class 
 Rate of progression from asymptomatic class to infected class 
 Natural death rate for females 
 Natural death rate for males 
 Death rate due to disease 
 Densitydependent reduction rate for females 
 Densitydependent reduction rate for males 
 Average harem size 
 The probability that the newborn of an asymptomatic mother is susceptible 
