Research Article

Complementary Variety: When Can Cooperation in Uncertain Environments Outperform Competitive Selection?

Table 1

Fitness landscapes in matrix form with types and environmental states and corresponding fitness values . The larger fitness value of the binary type split (horizontal) is denoted with capital letter and the less fit of the two with the small letter . Despite this simple case of two types, the covariance relation of (2) also holds also for more than two types. (a) schematizes a positive covariance between type fitness in environments and , while (b) schematizes a negative covariance, with superior type fitness along the diagonal of the matrix.
(a)

Population type level
Environ. state


(b)

Population type level
Environ. state