Identification of Fine-Scale Marine Benthic Ecoclines by Multiple Parallel Ordination

Table 4

Frequency and average abundance in presence sites per species as a function of affiliation to broad-scaled features.

	Frequency						Average abundance in presence sites						freq	occ	ab.avr	ab.tot
	C	SS	SF	D	DS	DF	C	SS	SF	D	DS	DF
Polychaeta
Pherusa falcata	1.00						1.67						0.11	3	1.67	5
Eumida sanguinea	0.67	0.17					2.50	1.00					0.11	3	2.00	6
Eunice pennata	0.67	0.17					2.50	2.00					0.11	3	2.33	7
Streblosoma bairdi	0.33	0.50	0.80	0.40	0.50		1.00	2.33	1.50	2.00	1.50		0.43	12	1.75	21
Ophelina acuminata	0.33	0.33	0.20	0.40	0.25		2.00	2.00	1.00	1.00	1.00		0.25	7	1.43	10
Maldane sarsi	0.33	0.17	0.60	0.40	0.25		9.00	5.00	3.67	8.50	3.00		0.29	8	5.62	45
Anobothrus gracilis	1.00	1.00	1.00	1.00	1.00	1.00	51.33	18.67	7.60	6.40	6.50	3.80	1.00	28	13.61	381
Glycera alba	1.00	1.00	1.00	1.00	1.00	1.00	10.00	8.83	10.60	11.20	8.25	6.20	1.00	28	9.14	256
Goniada maculata	1.00	1.00	1.00	1.00	1.00	1.00	17.67	11.00	12.00	14.00	12.75	10.60	1.00	28	12.61	353
Scoletoma fragilis	1.00	1.00	1.00	1.00	1.00	1.00	11.67	11.50	14.60	17.40	10.25	12.20	1.00	28	13.07	366
Spiophanes kroyeri	1.00	1.00	1.00	1.00	1.00	1.00	35.33	47.00	33.60	141.40	94.25	149.80	1.00	28	85.32	2389
Typosyllis cornuta	1.00	1.00	1.00	1.00	1.00	1.00	13.33	7.83	11.40	10.00	7.00	8.20	1.00	28	9.39	263
Pholoe inornata	1.00	1.00	1.00	1.00	1.00	0.80	5.67	2.50	4.00	3.00	2.25	3.75	0.96	27	3.37	91
Polycirrus norvegicus	1.00	1.00	1.00	1.00	1.00	0.80	6.00	7.67	7.80	9.20	2.00	5.50	0.96	27	6.63	179
Chaetozone setosa	1.00	1.00	1.00	1.00	1.00	0.60	28.00	8.33	10.60	12.80	53.00	23.67	0.93	26	20.54	534
Pista cristata	1.00	1.00	1.00	1.00	0.75	1.00	14.00	12.83	13.60	12.40	9.67	9.40	0.96	27	12.04	325
Polyphysia crassa	1.00	1.00	1.00	1.00	0.75	0.80	23.33	16.83	7.80	6.40	4.00	8.75	0.93	26	11.12	289
Heteromastus filiformis	1.00	1.00	0.80	1.00	1.00	1.00	19.33	5.67	7.50	46.40	44.00	11.00	0.96	27	21.67	585
Terebellides stroemi	1.00	1.00	0.80	1.00	1.00	1.00	9.33	8.83	9.00	6.20	10.00	4.60	0.96	27	7.81	211
Pseudopolydora paucibranchiata	1.00	1.00	0.40	0.40	0.75	0.60	194.33	9.50	30.50	6.00	6.33	4.67	0.68	19	39.26	746
Brada villosa	1.00	1.00	0.40	0.40	0.25	0.40	1.33	4.50	5.00	2.00	3.00	4.00	0.57	16	3.50	56
Prionospio cirrifera	1.00	0.83	0.60	1.00	1.00	1.00	60.67	7.00	21.67	14.60	10.50	8.20	0.89	25	17.52	438
Trichobranchus roseus	1.00	0.83	0.60	0.60	0.75	0.80	1.67	3.40	3.67	3.33	1.67	3.50	0.75	21	2.95	62
Prionospio fallax	1.00	0.67	1.00	0.80	1.00	0.80	25.33	16.75	8.00	4.25	11.00	5.00	0.86	24	11.00	264
Diplocirrus glaucus	1.00	0.67	0.80	1.00	1.00	0.60	3.67	3.75	4.25	3.60	13.50	4.00	0.82	23	5.52	127
Polycirrus medusa	1.00	0.50	0.40	0.60	0.50	0.60	2.67	1.67	3.50	1.00	12.00	1.33	0.57	16	3.19	51
Melinna elisabethae	1.00	0.50	0.20	0.60	0.50	0.60	6.67	5.67	1.00	1.67	1.00	3.00	0.54	15	3.60	54
Spiochaetopterus typicus	0.67	1.00	1.00	1.00	1.00	1.00	1.00	4.00	6.60	3.60	7.25	7.20	0.96	27	5.26	142
Aphrodita aculeata	0.67	0.67	0.40	0.20	0.50	0.40	3.00	1.75	1.00	4.00	1.00	1.50	0.46	13	1.85	24
Melinna cristata	0.67	0.50	0.40	0.80	1.00	1.00	2.50	2.67	3.00	4.25	2.75	3.80	0.71	20	3.30	66
Syllidia armata	0.67	0.33	0.80	0.60	0.25	0.60	2.50	1.50	1.25	2.00	1.00	1.33	0.54	15	1.60	24
Scalibregma inflatum	0.67	0.33	0.40	0.60	0.50	0.40	1.50	1.00	1.00	1.00	1.00	1.00	0.46	13	1.08	14
Amaena trilobata	0.67	0.17	0.40	0.40	0.50	0.80	2.00	2.00	1.00	5.00	1.00	1.50	0.46	13	2.00	26
Trichobranchus glacialis	0.33	0.67	0.20	0.40	1.00	0.60	1.00	2.25	5.00	3.00	1.25	1.33	0.54	15	2.00	30
Pista lornensis	0.33	0.17	0.60	0.40	0.75	0.80	1.00	1.00	1.67	1.50	1.00	1.25	0.50	14	1.29	18
Sige fusigera	0.33	0.17	0.20	0.40	0.25	0.40	1.00	1.00	3.00	2.50	1.00	1.00	0.29	8	1.62	13
Eupolymnia nebulosa	0.67	0.33	0.20	0.20		0.20	3.50	2.00	2.00	1.00		1.00	0.25	7	2.14	15
Glycera lapidum	1.00	0.33		0.40	0.25	0.20	11.67	6.50		1.50	1.00	1.00	0.32	9	5.89	53
Cossura longocirrata	0.67	0.17		0.20	0.25		1.50	1.00		1.00	1.00		0.18	5	1.20	6
Sphaerodorum gracilis	0.67	0.33		0.20			2.50	1.00		1.00			0.18	5	1.60	8
Pholoe assimilis	0.33	0.17		0.20			4.00	1.00		1.00			0.11	3	2.00	6
Chaetoparia nilssoni	0.33			0.60	0.50		1.00			1.00	1.00		0.21	6	1.00	6
Scoloplos armiger	0.67			0.20			19.00			3.00			0.11	3	13.67	41
Scoletoma cf. magnidentata		0.33	0.40	0.40	0.50	0.20		1.00	1.00	1.00	1.00	1.00	0.32	9	1.00	9
Amphicteis gunneri		0.17	0.40	0.20	0.25	0.40		2.00	1.00	1.00	2.00	2.00	0.25	7	1.57	11
Ophiodromus flexuosus		0.33	0.20	0.20	0.25			1.50	1.00	2.00	1.00		0.18	5	1.40	7
Phyllodoce rosea		0.33	0.40		0.50	0.40		1.50	1.00		2.00	1.50	0.29	8	1.50	12
Phyllodoce groenlandica		0.33		0.20		0.20		1.00		1.00		1.00	0.14	4	1.00	4
Dipolydora cf. coeca		0.17						11.00					0.04	1	11.00	11
Petaloproctus borealis		0.17						5.00					0.04	1	5.00	5
Sabellides octocirrata			0.20	0.40	0.25	0.20			1.00	1.00	1.00	1.00	0.18	5	1.00	5
Orbinia sertulata			0.20	0.40	0.25				2.00	1.00	1.00		0.14	4	1.25	5
Nephtys ciliata			0.20		0.50	0.40			1.00		2.00	1.00	0.18	5	1.40	7
Glycera unicornis			0.60		0.50				1.00		1.00		0.18	5	1.00	5
Rhodine loveni				0.20						5.00			0.04	1	5.00	5
Mollusca
Hiatella arctica	0.67	0.33					4.00	2.00					0.14	4	3.00	12
Lepeta caeca	0.67	0.17					20.00	6.00					0.11	3	15.33	46
Mya truncata	0.67	0.17					1.50	1.00					0.11	3	1.33	4
Arctica islandica	0.33	0.50	0.20				1.00	1.67	1.00				0.18	5	1.40	7
Thyasira equalis	1.00	1.00	1.00	1.00	0.75	1.00	9.00	14.67	12.20	17.20	37.00	18.40	0.96	27	17.22	465
Astarte montagui	1.00	1.00	0.80	0.80	0.75	0.60	13.00	5.00	6.25	5.25	6.33	3.00	0.82	23	6.22	143
Caudofoveata	1.00	0.83	0.60	0.80	0.50	0.40	1.67	1.80	2.00	1.75	1.50	2.00	0.68	19	1.79	34
Ennucula tenuis	1.00	0.67	1.00	1.00	1.00	1.00	39.00	68.00	72.40	106.80	130.00	82.20	0.93	26	85.23	2216
Pseudamussium peslutrae	1.00	0.50	0.40	0.20	0.25	0.20	2.67	2.00	1.00	1.00	1.00	1.00	0.39	11	1.73	19
Thyasira sarsi	0.67	0.83	0.80	1.00	1.00	1.00	3.50	3.20	6.75	10.00	11.25	12.40	0.89	25	8.28	207
Abra nitida	0.33	0.50	0.40	0.80	0.50	1.00	5.00	2.67	2.00	3.00	5.00	2.60	0.61	17	3.06	52
Thyasira flexuosa	0.33	0.33	0.80	1.00	1.00	1.00	3.00	7.00	5.50	11.40	11.50	12.80	0.75	21	9.81	206
Philine scabra	0.33	0.33	0.20	0.20	0.25	0.20	2.00	1.00	1.00	2.00	2.00	1.00	0.25	7	1.43	10
Corbula gibba	1.00	0.83	0.20			0.40	7.00	20.60	5.00			1.50	0.39	11	12.00	132
Parvicardium minimum	0.67	0.17				0.20	1.00	1.00				1.00	0.14	4	1.00	4
Macoma calcarea	0.67		0.40	0.40	0.50	0.60	7.00		6.00	2.00	7.00	8.67	0.39	11	6.36	70
Astarte elliptica	0.67		0.20	0.60		0.20	3.00		2.00	3.33		1.00	0.25	7	2.71	19
Thyasira gouldi		0.33	0.20	0.40		0.80		1.00	1.00	3.00		2.00	0.32	9	1.89	17
Nuculana pernula		0.33			0.25			1.00			2.00		0.11	3	1.33	4
Buccinum undatum			0.40	0.40	0.25	0.40			1.00	1.00	1.00	1.00	0.25	7	1.00	7
Echinodermata
Strongylocentrotus droebachiensis	0.67						2.00						0.07	2	2.00	4
Amphiura chiajei	0.67	0.83	0.40				1.50	3.60	3.50				0.32	9	3.11	28
Ophiocten affinis	1.00	1.00	0.60	0.60	0.50	0.20	13.33	6.17	4.33	3.67	1.00	2.00	0.64	18	5.83	105
Ophiura albida	1.00	0.50	0.40	0.40	0.50	0.40	2.00	1.33	1.00	1.00	2.50	2.00	0.50	14	1.64	23
Amphiura filiformis	0.67	0.50	0.60	0.40	0.25	0.40	1.50	5.67	3.00	1.00	7.00	2.00	0.46	13	3.23	42
Ophiura robusta	0.67	0.33		0.20			2.00	1.00		1.00			0.18	5	1.40	7
Amphipholis squamata	0.33	0.67	0.40	0.20		0.20	2.00	1.25	1.00	4.00		2.00	0.32	9	1.67	15
Brissopsis lyrifera	0.67		0.40			0.20	1.00		1.00			1.00	0.18	5	1.00	5
Echinocardium flavescens		0.33	0.40	0.40	0.25	0.60		1.00	1.50	1.00	1.00	1.67	0.36	10	1.30	13
Polychaeta
Euchone rubrocinta	0.33						1.00						0.04	1	1.00	1
Exogone verugera	0.33						1.00						0.04	1	1.00	1
Harmothoe cf. fragilis	0.33						2.00						0.04	1	2.00	2
Laonice cirrata	0.33						2.00						0.04	1	2.00	2
Nicolea zostericola	0.33						1.00						0.04	1	1.00	1
Pherusa plumosa	0.33						2.00						0.04	1	2.00	2
Pholoe pallida	0.33						1.00						0.04	1	1.00	1
Praxillura longissima	0.33						1.00						0.04	1	1.00	1
Rhoine gracilior	0.33						2.00						0.04	1	2.00	2
Gyptis rosea	0.67			0.20			1.00			1.00			0.11	3	1.00	3
Eteone cf. longa	0.67				0.25		1.00				1.00		0.11	3	1.00	3
Galathowenia oculata		0.17						1.00					0.04	1	1.00	1
Laetmonice filicornis		0.17						1.00					0.04	1	1.00	1
Orbinia norvegica		0.17						1.00					0.04	1	1.00	1
Typosyllis armillaris		0.17						1.00					0.04	1	1.00	1
Parougia eliasoni				0.20	0.25	0.20				1.00	1.00	1.00	0.11	3	1.00	3
Flabelligera affinis				0.20						1.00			0.04	1	1.00	1
Paraamphinome jeffreysii					0.25	0.20					2.00	1.00	0.07	2	1.50	3
Amage auricola					0.25						2.00		0.04	1	2.00	2
Mediomastus fragilis					0.25						1.00		0.04	1	1.00	1
Neoamphitrite grayi						0.20						1.00	0.04	1	1.00	1
Mollusca
Parvicardium pinnulatum	0.33						1.00						0.04	1	1.00	1
Tellimya ferruginosa	0.67	0.17					1.00	1.00					0.11	3	1.00	3
Aporrhais pespelecani	0.33	0.17	0.20				1.00	1.00	1.00				0.11	3	1.00	3
Cuspidaria cuspidata		0.17						1.00					0.04	1	1.00	1
Kurtiella bidentata		0.17						1.00					0.04	1	1.00	1
Abra alba		0.17				0.20		1.00				1.00	0.07	2	1.00	2
Astarte sulcata			0.20	0.20		0.20			1.00	1.00		1.00	0.11	3	1.00	3
Antalis entalis			0.20						1.00				0.04	1	1.00	1
Diaphana minuta			0.20						1.00				0.04	1	1.00	1
Cylichna alba					0.25						1.00		0.04	1	1.00	1
Lunatia montagui						0.20						1.00	0.04	1	1.00	1
Mytilus edulis						0.20						2.00	0.04	1	2.00	2
Rissoidae						0.20						2.00	0.04	1	2.00	2
Echinodermata
Echinocardium cordatum		0.17			0.25			2.00			1.00		0.07	2	1.50	3
Ophiura ophiura		0.17			0.25			1.00			1.00		0.07	2	1.00	2
Ophiopholis aculeata				0.20						2.00			0.04	1	2.00	2
Ophiothrix fragilis						0.20						1.00	0.04	1	1.00	1

Freq: total site frequency; occ: total site occurrences; ab.avr: overall average abundance in presence sites; ab.tot: overall total abundance in presence sites.