Optimization of the Spatiotemporal Parameters in a Dynamical Marine Ecosystem Model Based on the Adjoint Assimilation

Table 1

Ecological parameters and their initial values in the model.

Symbol	Parameter	Value	Unit
𝑉𝑚	Maximum uptake rate of nutrient by phytoplankton	0.8	Day^−1
𝐾𝑠	Half-saturation constant for nutrient uptake	0.25	(N) mmol · m^−3
𝐴𝑄10	Temperature coefficient for phytoplankton growth at 10°C	2.08	—
𝑊𝑃	Sinking velocity of phytoplankton	1.0	m · day^−1
𝐺𝑚	Zooplankton maximum grazing rate	0.50	day^−1
𝜆	Zooplankton Ivlev constant	0.20	(N)
𝐵𝑄10	Temperature coefficient for zooplankton growth; at 10°C	3.10	—
𝐷𝑧	Zooplankton mortality rate	0.05	Day^−1
𝛾	Assimilation ratio for zooplankton	0.30	—
𝐷𝑝	Phytoplankton mortality rate	0.10	Day^−1
𝑒	Remineralization rate of detritus	0.0212	Day^−1
𝜃	Excretion ratio for zooplankton	0.40	—
𝑊𝑑	Sinking velocity of detritus	1.00	m · day^−1