Translational Approach to Behavioral Learning: Lessons from Cerebellar Plasticity

Figure 4

Directional learning during pursuit eye movements. (a) Representative eye movement during pursuit of a target moving first toward the right and then in oblique upward direction, the inset indicates the target motion in polar coordinates. (b) Eye velocity trace of the movement illustrated in (a). Gray and black traces indicate data from representative trials before learning and after at least 100 learning trials, respectively. The dashed traces indicate the velocity step signal of the target motion. Note that the vertical step (lower part) delayed the horizontal one (upper part) by 250 ms. The arrow pointing down and right indicates the learned response. The arrow pointing up and left indicates the hardwired visual response to the change in target direction, (adapted from [83] with permission).