Effects of Habitat and Human Activities on Species Richness and Assemblages of Staphylinidae (Coleoptera) in the Baltic Sea Coast

Table 2

Mean values of environmental parameters for the differentiated staphylinid assemblages; different exponents indicate significant differences by ANOVA and consecutive LSD test.

Parameter	Wooded cliffs	Woodless cliffs	Lagoon	Sandy beach	Primary dune	Shingle beach
Soil moisture (%)	^a18±3.0	^b8±1.1	^a31±5.9	^c1±0.5	^c1±0.5	^c1±0.6
Wood	yes	no	no	no	no	no
Sand content (%)	^b36±6	^c46±3	^c51±16	^a89±9	^a92±9	^b22±21
Shingle content (%)	^b5±2	^b8±2	^b21±20	^b8±10	^b6±8	^a77±20
Soil pH	^b7.4±0.1	^a7.9±0.2	^b7.5±0.2	^b7.3±0.2	^c7.2±0.1	^a7.9±0.3
Organic content of soil (%)	^a6.6±2.6	^b1.1±0.1	^a5.6±0.9	^b0.2±0.2	^b1.4±2.2	^b0.3±0.1