|
(1) Antenna with 12 flagellomeres | 2 |
Antenna with 13 flagellomeres | 6 |
(2) Apical lamellae of median volsella bulbous, Oriental China | bullus Wang & Guo |
Apical lamellae of median volsella not bulbous | 3 |
(3) Apex of anal point broad, India, Malaysia | madarihatensis Kyerematen, Andersen & Sæther |
Apex of anal point narrow | 4 |
(4) Anal tergite bands nearly horizontal, Oriental China | polychaetus Wang & Guo |
Anal tergite bands V-shaped | 5 |
(5) Anal tergite with narrowly interrupted basal band, Malaysia | kuantanensis Kyerematen, Andersen & Sæther |
Anal tergite without basal band, India | nudicornus n. sp. |
(6) Median volsella long, reaching beyond apex of inferior volsella | 7 |
Median volsella short, not reaching beyond apex of inferior volsella | 10 |
(7) Median volsella apically with two distinct plates, Indonesia | adjectus (Johannsen) |
Median volsella with distal lamellate apical setae never fused into plate | 8 |
(8) Anal tergite band V-shaped, Thailand, Australia | oss Cranston |
Anal tergite band transverse | 9 |
(9) Anal tergite band separate; median volsella conspicuously slender, Thailand | minusculus Kyerematen, Andersen & Sæther |
Anal tergite band medially fused; median volsella not slender, Oriental China, Ghana | buculicaudus Kyerematen in Kyerematen, Andersen & Sæther |
(10) Posterior margin of tergite IX with projections or shoulders to each side | 11 |
Posterior margin of tergite IX triangular, rounded, or at most straight | 13 |
(11) Tergite IX posterior margin with well-developed projections; gonostylus abruptly tapered | 12 |
Tergite IX posterior margin without such projections, but with distinct shoulders, gonostylus not abruptly tapered, Indonesia, Australia | trivittatus (Johannsen) |
(12) Inferior volsella hand shower shaped; basal anal tergite band joined medially, India | caputimberus n. sp. |
Inferior volsella not as above, basal anal tergite band interrupted, Indonesia | additus (Johannsen) |
(13) Digitus well developed, extending beyond margin of superior volsella | 14 |
Digitus small or absent, not extending beyond margin of superior volsella | 21 |
(14) Median volsella not reaching apex of superior volsella; if gonostylus abruptly tapering distally, then apical portion parallel-sided and straight | 15 |
Median volsella at least reaching apex of superior volsella; if gonostylus abruptly tapered, then apical portion not parallel-sided and straight | 20 |
(15) Superior volsella knob- or hook-like posterior extension | 16 |
Superior volsella rounded, ovoid, oblong or thumb-like | 18 |
(16) Superior volsella with posterior extension knob-like, Thailand | orientalis Moubayed |
Superior volsella with posterior extension hook-like | 17 |
(17) Superior volsella greatly hooked; apical setae of superior volsella not fused into plate; Thailand, Lebanon, Europe | reissi Lehmann |
Superior volsella slightly hooked; apical setae of superior volsella fused into plate; Oriental and Palaearctic China, Europe, North Africa and Canada | muscicola Thienemann |
(18) Anal point tapering; Oriental Japan | amamiflavus Sasa |
Anal point spatulate | 19 |
(19) Superior volsella rectangular with rounded margin; basal tergite band medially joined; Oriental China, Palaearctic Japan | tamatertius Sasa |
Superior volsella rounded; basal tergite band absent; Oriental Japan | okisimplex Sasa |
(20) Gonostylus abruptly tapered with apical portion curved; superior volsella with large, bluntly rounded apical projection; median volsella apparently without apical plate, India, Indonesia, Oriental China | acerbus Johannsen |
Gonostylus not abruptly tapered; superior volsella subquandrangular; median volsella with wide apical plate; Palaearctic Japan, Oriental and Palaearctic China | tamaquartus Sasa |
(21) Abdomen banded; anal point crests long, proximally fused forming an arc; superior volsella rounded; median volsella without plate | 22 |
Abdomen not banded; hypopygium not with above configuration | 23 |
(22) Gonostylus not abruptly tapered plate, India, Indonesia, Oriental and Palaearctic Japan, Palaearctic China | aestuarius (Tokunaga) |
Gonostylus abruptly tapered, Oriental China | liuae Wang & Guo |
(23) Gonostylus more or less abruptly tapered, with distinctly parallel-sided apical portion or with curved apex; superior volsella with pronounced posterior extension; median volsella extending beyond apex of superior volsella | 24 |
If gonostylus abruptly tapered, then without parallel-sided or curved apex | 25 |
(24) Gonostylus with apical portion distinctly parallel-sided; superior volsella with posterior extension long, digitiform; Malaysia | phaselus Kyerematen, Andersen & Sæther |
Gonostylus with apex curved; superior volsella with posterior extension broad and rounded; Thailand | thailandensis Moubayed |
(25) Anal tergite bands V-shaped and medially joined | 26 |
If anal tergite bands V-shaped, then not medially joined | 27 |
(26) Anal point nonspatulate; median volsella extremely beyond apex of superior volsella; AR > 0.60, Indonesia | tobaseptidecimus Kikuchi & Sasa |
Anal point spatulate; median volsella not extending beyond apex of superior volsella; AR < 0.40, India | spinicornus n. sp. |
(27) Apex of anal point spatulate; gonostylus tapering abruptly or gradually | 28 |
Apex of anal point parallel-sided or tapering; gonostylus tapering gradually | 34 |
(28) Superior volsella with pronounced, hook-like posterior extension, Thailand | falcipedius Kyerematen, Andersen & Sæther |
Superior volsella rounded or rectangular | 29 |
(29) Superior volsella rectangular, Oriental China | quadratus Wang & Guo |
Superior volsella rounded | 30 |
(30) Gonostylus tapering gradually | 31 |
Gonostylus abruptly tapered in apical portion | 33 |
(31) Anal point crests proximally fused, forming an arc, Thailand | beccus Kyerematen, Andersen & Sæther |
Anal point crest V-shaped | 32 |
(32) Median volsella reaching beyond apex of superior volsella; basal anal tergite bands absent; Thailand, Lebanon, Europe | curtistylus (Goetghebuer) |
Median volsella short, not reaching apex of superior volsella; basal anal tergite bands present, Thailand | falcatus Kyerematen, Andersen & Sæther |
(33) Median volsella recurved; AR about 0.70, Thailand | sessilipersonatus Kyerematen, Andersen & Sæther |
Median volsella not markedly recurved; AR about 0.40, Thailand | soelii Kyerematen, Andersen & Sæther |
(34) Median volsella extending beyond apex of superior volsella; anal point crests proximally fused, forming an arc, Thailand | pallidus Kyerematen, Andersen & Sæther |
Median volsella not reaching beyond apex of superior volsella; anal point crests V-shaped | 35 |
(35) Anal point broad, parallel–sided; AR about 0.1–0.3, Thailand | koraensis Kyerematen, Andersen & Sæther |
Anal point lanceolate; AR about 0.3–0.4, Thailand | verticillus Kyerematen, Andersen & Sæther |
|