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| (1) Tergite VIII with caudolateral comb; T II–V with anterior paired patches of spinules; thoracic horn with knee-like bend, heavily sclerotized | 2 |
| Tergite VIII with single spur; T II–IV, II–V or II–VI with paired patches of spinules; thoracic horn may be sharply bent but not knee-like | 3 |
| (2) Hook row of about 90 hooklets; anal lobe fringe of much less than 20 taeniae | additus (Johannsen) |
| Hook row of 60–70 hooklets; anal lobe fringe of about 20 taeniae | trivittatus (Johannsen) |
| (3) Thoracic horn sharply bent at midlength; T II–IV with oral, paired, rounded patches of spinules | 4 |
| Thoracic horn not sharply bent; T II–V or II–VI with spinule patches | 5 |
| (4) Tergite VIII with 3 lateral taeniae | thailandensis Moubayed |
| Tergite VIII with 5 lateral taeniae | oss Cranston |
| (5) Tergites II–VI with sharply defined, paired spinule patches | 6 |
| Tergites II–V with sharply defined patches of spinules | 7 |
| (6) Anal lobe with one large dorsal seta; thoracic horn with few fine spinules | curtistylus (Goetghebuer) |
| Anal lobe with two short dorsal setae; thoracic horn with many fine spinules in distal half | orientalis Moubayed |
| (7) Anal lobe without dorsal setae; distal half of thoracic horn with many spinules | adjectus (Johannsen) |
| Anal lobe with long dorsal setae; distal half of thoracic horn with or without any spinules | 8 |
| (8) Distal half of thoracic horn without any spinules | nudicornus n. sp. |
| Distal half of thoracic horn with few to many spinules | 9 |
| (9) Tergites II and III with transversely elongated or rectangular spine patches | reissi Lehmann |
| Tergites II-III with circular or elliptical spine patches | 10 |
| (10) Numerous spinules on distal 2/3 of the thoracic horn | spinicornus n. sp. |
| Spinules few on distal 1/3 of the thoracic horn | tamaquartus Sasa |
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